Skip to main content
The University of Chicago Marine Biological Laboratory
Menu
expanded
collapsed
Weather
VISIT THE MBL
Utility
Contact
Careers
Give
About
Back
About the MBL
Leadership
Back
Leadership - Overview
Board of Trustees Committees
Nipam Patel, Director
Back
Nipam Patel, Director - Overview
Gallery of Nipam Patel Research Images
Courtney Buckley
Alison Crawford
Samantha Cummis
Mary S. Harrington
Linda Hyman
Scott Koerner
Marie Russell
Sharon Stanczak
Anne W. Sylvester
Jennifer Walton
MBL Director History
Advisory Bodies
Advisory Bodies
Back
Advisory Bodies - Overview
MBL Society
Back
MBL Society - Overview
Anne Hart
MBL Society Members
MBL Society Membership Criteria and Procedures
MBL Community Advisory Council Charter
MBL Science Council Charter
MBL Society Steering Committee
MBL Community
MBL Community
Back
MBL Community - Overview
Land Acknowledgement Statement
MBL Community Committee
University of Chicago
Financial Statements
Careers & Internships
Careers & Internships
Back
Careers & Internships - Overview
Our Community
Benefits Summary
Equal Employment Opportunity
Back
Equal Employment Opportunity - Overview
Accommodation of Individuals with Disabilities
Drug-Free Workplace
Harassment-Free Workplace
Notice of Nondiscrimination
Staff Toolbox
Electronic Posters
Contact HR
History & Archives
History & Archives
Back
History & Archives - Overview
Nobel Laureates Affiliated with the MBL
Legacy of Leadership
Back
Legacy of Leadership - Overview
Cornelia Clapp
Jewel Plummer Cobb
Jean Clark Dan and Katsuma Dan
Ernest Everett Just
Stephen W. Kuffler
Joseph L. Martinez, Jr. and James G. Townsel
Lilian Vaughan and Thomas Hunt Morgan
Seeing Life Exhibit
Back
Seeing Life Exhibit - Overview
Seeing Life at MBL
Bronze Memorial Tablet
Charles Otis Whitman
Edmund Beecher Wilson
Thomas Hunt Morgan
Cornelia Clapp
Ernest Everett Just
Ethel Browne Harvey
Albert Szent-Györgyi
Jean Clark Dan
Katsuma Dan
"Last One to Go"
Shinya Inoué
Polarized Light Microscopy
Seeing Spindles
Video Microscopy
Advertisements from the Collecting Net
Exhibit Videos
Alumni
Alumni
Back
Alumni - Overview
ARTC Reunions
Back
ARTC Reunions - Overview
Embryology 130th Anniversary Celebration
SES 25th Anniversary Celebration and SBD Welcome Symposium
Neurobiology 50th Anniversary Symposium
Microbial Diversity 50th Anniversary Symposium
Class Photos
Back
Class Photos - Overview
Biology of Aging Class Photos
Back
Biology of Aging Class Photos - Overview
Biology of Aging 2020s
Gene Regulatory Networks for Development Class Photos
Back
Gene Regulatory Networks for Development Class Photos - Overview
Gene Regulatory Networks for Development 2000s
Gene Regulatory Networks for Development 2010s
Biology of Parasitism Class Photos
Back
Biology of Parasitism Class Photos - Overview
Biology of Parasitism 1980s
Biology of Parasitism 1990s
Biology of Parasitism 2000s
Biology of Parasitism 2010s
Biology of Parasitism 2020s
Brains, Minds and Machines Class Photos
Back
Brains, Minds and Machines Class Photos - Overview
Brains, Minds and Machines 2010s
Brains, Minds and Machines 2020s
Embryology Class Photos
Back
Embryology Class Photos - Overview
Embryology 1880s
Embryology 1940s
Embryology 1950s
Embryology 1960s
Embryology 1970s
Embryology 1980s
Embryology 1990s
Embryology 2000s
Embryology 2010s
Embryology 2020s
Endocrine-Disrupting Chemicals: Hazards and Opportunities Class Photos
Back
Endocrine-Disrupting Chemicals: Hazards and Opportunities Class Photos - Overview
Endocrine-Disrupting Chemicals: Hazards and Opportunities 2020s
Frontiers in Reproduction Class Photos
Back
Frontiers in Reproduction Class Photos - Overview
Frontiers in Reproduction 1990s
Frontiers in Reproduction 2000s
Frontiers in Reproduction 2010s
Frontiers in Reproduction 2020s
Frontiers in Stem Cells and Regeneration Class Photos
Back
Frontiers in Stem Cells and Regeneration Class Photos - Overview
Frontiers in Stem Cells and Regeneration 2010s
Frontiers in Stem Cells and Regeneration 2020s
History of Biology Class Photos
Back
History of Biology Class Photos - Overview
History of Biology 1980s
History of Biology 1990s
History of Biology 2000s
History of Biology 2010s
History of Biology 2020s
Marine Ecology Class Photos
Back
Marine Ecology Class Photos - Overview
Marine Ecology 1960s
Marine Ecology 1970s
Marine Ecology 1980s
Marine Ecology 1990s
Medical Informatics Class Photos
Back
Medical Informatics Class Photos - Overview
Medical Informatics 1990s
Medical Informatics 2000s
Microbial Diversity Class Photos
Back
Microbial Diversity Class Photos - Overview
Microbial Diversity 1970s
Microbial Diversity 1980s
Microbial Diversity 1990s
Microbial Diversity 2000s
Microbial Diversity 2010s
Molecular and Cell Biology of Symbiosis Class Photos
Back
Molecular and Cell Biology of Symbiosis Class Photos - Overview
Molecular and Cell Biology of Symbiosis 2020s
Molecular Evolution Class Photos
Back
Molecular Evolution Class Photos - Overview
Molecular Evolution 2000s
Molecular Evolution 2010s
Molecular Mycology Class Photos
Back
Molecular Mycology Class Photos - Overview
Molecular Mycology 1990s
Molecular Mycology 2000s
Molecular Mycology 2010s
Molecular Mycology 2020s
Neural Systems & Behavior Class Photos
Back
Neural Systems & Behavior Class Photos - Overview
Neural Systems & Behavior 1970s
Neural Systems & Behavior 1980s
Neural Systems & Behavior 1990s
Neural Systems & Behavior 2000s
Neural Systems & Behavior 2010s
Neural Systems & Behavior 2020s
Neurobiology Class Photos
Back
Neurobiology Class Photos - Overview
Neurobiology 1970s
Neurobiology 1980s
Neurobiology 1990s
Neurobiology 2000s
Neurobiology 2010s
Neurobiology 2020s
Physiology Class Photos
Back
Physiology Class Photos - Overview
Physiology 1890s
Physiology 1950s
Physiology 1960s
Physiology 1970s
Physiology 1980s
Physiology 1990s
Physiology 2000s
Physiology 2010s
Physiology 2020s
Semester in Environmental Science Class Photos
Back
Semester in Environmental Science Class Photos - Overview
SES 2000s
SES 2010s
SES 2020s
Summer Program in Neuroscience, Excellence and Success Class Photos
Back
Summer Program in Neuroscience, Excellence and Success Class Photos - Overview
SPINES 2020s
Zebrafish Class Photos
Back
Zebrafish Class Photos - Overview
Zebrafish 2020s
Support the Alumni Fund
Contact Alumni Staff
Campus Facilities & Resources
Campus Facilities & Resources
Back
Campus Facilities & Resources - Overview
Housing
Back
Housing - Overview
Apply for Housing
Campus Info for Guests
Reservation Process
General Information and Rates
Housing Services
Housing Policies
Housing Buildings
Back
Housing Buildings - Overview
Brick Dorms and Apartments
Memorial Circle Cottages
Devil's Lane Cottages
David House
Drew House
Smith Cottage
11 North Street
Ebert Hall
Swope Building
Crane House
Veeder House
Local Rentals
Contact Housing Staff
Dining
Back
Dining - Overview
Dining Venues
Dietary Accommodations
Swope Café Weekly Specials
Dining Programs
MBL Dining Hall Meals
Sodexo Sustainability @ MBL
Contact Dining Staff
Campus Maps
Parking, Transportation and Grounds
Back
Parking, Transportation and Grounds - Overview
Day Visitor Parking Request
MBL Shuttle Service
Back
MBL Shuttle Service - Overview
MBL Policy For Shuttle Van Use
Contact Transportation & Grounds Staff
Information Technology
Back
Information Technology - Overview
Multimedia Services
Telecommunications
Shipping and Receiving
Security and Campus Emergencies
Supplier User Guide
Summer Camp & Children's Programs
Bicycle Rentals
Boating Club
Campus Accessibility
Single User Restrooms
Local Area Information
Local Area Information
Back
Local Area Information - Overview
Travel Information
Local Hotels
Sculptures on the MBL Campus
Sustainability
Sustainability
Back
Sustainability - Overview
Recycling
Composting
Electric Vehicles
Nitrogen and Carbon Footprint Tracking
Water Kiosk and Bottle Refill Stations
Friends of the MBL/Get Involved
Production Studio
Campus Health
Research
Back
Groundbreaking Research at the MBL
Research Centers
Research Centers
Back
Research Centers - Overview
Ecosystems Center
Back
Ecosystems Center - Overview
About The Ecosystems Center
Ecosystems Center 50th Anniversary Celebration
Back
Ecosystems Center 50th Anniversary Celebration - Overview
50th Anniversary Program
Ecosystems Education Info
Employment/Internships
Research Projects
Back
Research Projects - Overview
Long-Term Ecological Research (LTER)
MBL Greenhouse
Oceanic Flux Program (OFP)
Back
Oceanic Flux Program (OFP) - Overview
OFP Collaborations & Outreach
Back
OFP Collaborations & Outreach - Overview
Agreement for the Distribution of Oceanic Flux Program Sediment Trap Material and Data
OFP Bibliography
Microscopic Photos
OFP Cruise Photos
OFP Staff
Contact Us
Terrestrial Isotope Biogeochemistry
Ecosystem Modeling
Back
Ecosystem Modeling - Overview
Frontiers Model
Hydra Model
General Ecosystem Model (GEM)
Back
General Ecosystem Model (GEM) - Overview
GEM Download Page
Multiple Element Limitation (MEL)
Back
Multiple Element Limitation (MEL) - Overview
MBL MEL IV: Eight resources
MBL MEL V: Annual model
MEL Download Page
MEL II: Two elemental resources
Modelshell
Non-Equilibrium, Stable-Isotope Simulator (NESIS)
Terrestrial Ecosystem Model (TEM)
Tropical Forest Regrowth (TFR)
Center Reports
Stable Isotope Laboratory
Back
Stable Isotope Laboratory - Overview
Our Equipment
Sample Requirements
Services & Charges
Shipping & Turnaround
Media from the Lab
Ecosystems Publications
Ecosystems Center Faculty & Staff
Back
Ecosystems Center Faculty & Staff - Overview
Paul A. Steudler
Eugene Bell Center
Back
Eugene Bell Center - Overview
About the Bell Center
Bell Center Faculty and Staff
Back
Bell Center Faculty and Staff - Overview
Hanlon Lab
Back
Hanlon Lab - Overview
About Hanlon Lab
Hanlon Book and Book Chapters
Hanlon Lab Course Material
Hanlon Lab in the News
Hanlon Photography
Hanlon Publications
Hanlon Research Areas
Hanlon Videos
Horb Lab
Back
Horb Lab - Overview
Horb Research
Kate MacCord
Oldenbourg Lab
Back
Oldenbourg Lab - Overview
Oldenbourg Gallery
Oldenbourg Publications
Rosenthal Lab
Back
Rosenthal Lab - Overview
High-level RNA editing in cephalopods
Marine Model Organism Development
Rosenthal Lab Members
Site-directed RNA editing
Structural and functional connectivity of squid chromatophores
Shribak Lab
Back
Shribak Lab - Overview
Shribak Gallery
Featured Projects
Back
Featured Projects - Overview
Seeing Cells Exhibit
Back
Seeing Cells Exhibit - Overview
Seeing Cells in Life
Theory of Cells as Basic Units of Life
Protoplasm as the Living Stuff
Cell Specialization
From Fertilization to Organisms
Presenting and Representing "The" Cell
Experimental Manipulations
Resolving the Inside of Cells
Inside Living Cells
Seeing How Cell Parts Move
Seeing Cell Aggregates
Designing Synthetic Cells
Imagining Cells
Seeing Cells Videos
Tropical Seaweed Farm Project
Back
Tropical Seaweed Farm Project - Overview
Our Research
Tropical Seaweed News
Back
Tropical Seaweed News - Overview
Loretta Roberson is Funded to Implement Seaweed Farms for Biomass Production
Meet our new project coordinator, Dr. Gretchen Grebe!
Members of the ANCIENT MARINER consortium successfully install 24,500 ft of line with young kelp sporophytes in Kodiak, AK
Our first research farm is in the water!
Part 1: Damisela’s Concept, Construction, and Cruise to Puerto Rico
The Nature Conservancy and the Belize Women’s Seaweed Farmers Association featured on The Fish Site
Tropical Seaweed Field Sites
Project Team
Photo Gallery
Press
Bell Center Publications
Back
Bell Center Publications - Overview
Bell Center Publications 2024
Bell Center Publications 2023
Bell Center Publications 2022
Bell Center Publications 2021
Contact Bell Center
Josephine Bay Paul Center
Back
Josephine Bay Paul Center - Overview
About the JPBC
Bay Paul Publications
Back
Bay Paul Publications - Overview
JBPC Publications 2024
JBPC Publications 2023
JBPC Publications 2022
JBPC Publications 2021
JBPC Publications: 2011 - 2020
JBPC Publications: 2000 - 2010
JBPC Publications: 1996 - 1999
The Keck Facility
Back
The Keck Facility - Overview
Illumina Marker Gene and Shotgun Sequencing
Illumina Sequencing Requirements
Mobile Genetic Element Cluster
2024 CGUE Conference on Unicellular Eukaryotes
Back
2024 CGUE Conference on Unicellular Eukaryotes - Overview
CGUE Registration and Travel Information
CGUE Speakers
CGUE Early Career Events and More
CGUE Program
Bay Paul Center Faculty and Staff
Whitman Center
Back
Whitman Center - Overview
About the Whitman Center
Registration and Housing
Whitman Directory
Back
Whitman Directory - Overview
2024 Whitman Fellows
2024 Whitman Scientists
Pre-arrival, Arrival & Departure
Back
Pre-arrival, Arrival & Departure - Overview
Getting Started
Labs, Equipment, and Facilities
Responsible Conduct
Shut Down and Winter Storage
Faculty Directory
Whitman Directory
Researchers and Support Staff
Strategic Initiatives
Strategic Initiatives
Back
Strategic Initiatives - Overview
Imaging Innovation Initiative
New Research Organisms
Research Organisms
Research Organisms
Back
Research Organisms - Overview
African Clawed Frog
Atlantic Horseshoe Crab
Axolotl
Bat Sea Star
California Two-Spot Octopus
Comb Jelly
Common Cuttlefish
Flamboyant Cuttlefish
Green Algae
Hummingbird Bobtail Squid
Lesser Pacific Striped Octopus
Little Skate
Longfin Squid
Northern Star Coral
Parhyale
Rotifer
Sea Lamprey
Sea Robin
Striped Bass
MBL/UChicago PhD Graduate Program
Scientific Publications
Scientific Publications
Back
Scientific Publications - Overview
2024 Scientific Publications
2023 Scientific Publications
Who is at the MBL
Back
Who is at the MBL - Overview
Adjunct Scientists
MBL Postdoctoral Association
Back
MBL Postdoctoral Association - Overview
2025 Postdoctoral Symposium
Back
2025 Postdoctoral Symposium - Overview
2025 Postdoctoral Symposium Schedule
PDA Events
PDA Volunteer Opportunities
MBLWHOI Library
Resources & Research Facilities
Resources & Research Facilities
Back
Resources & Research Facilities - Overview
Imaging Resources
Back
Imaging Resources - Overview
Microscopes in Development
Central Microscopy Facility
Back
Central Microscopy Facility - Overview
CMF Equipment
Scheduling Time and Trainings
MLSC Imaging Community
Contact CMF
Laboratory Operations
Back
Laboratory Operations - Overview
Reserve Equipment
Back
Reserve Equipment - Overview
Balances
Microscopy
General Use Rooms
Submersible Heaters
Stockroom Information
Marine Resources Center
Back
Marine Resources Center - Overview
About the MRC
MRC Catalog
MRC Services
Small Boat Program
Research at the MRC
Cephalopods
Back
Cephalopods - Overview
Cephalopod Breeding Center
Order Organisms & Services
Back
Order Organisms & Services - Overview
Ordering Vertebrates
Tank Space Form
Gemma Tracker
Contact MRC Staff
The Microbiome Center
Zebrafish Facility
Back
Zebrafish Facility - Overview
Zebrafish Use
The National Xenopus Resource
Back
The National Xenopus Resource - Overview
About the NXR
Ordering and Pricing
Protocols
Back
Protocols - Overview
NXR BL-7 Vapor Shipper
NXR Cryosperm IVF (Straws)
NXR Cryosperm IVF (Tubes)
NXR Dry Ice Shipment Arrival
NXR Slow-Freezing Protocol
NXR Sperm Cryopreservation Protocol (X. laevis)
3D Printing Files
Mutant Frog Stocks
Transgenic Tropicalis
Transgenic Laevis
Back
Transgenic Laevis - Overview
Xla.Tg (CMV:Brainbow-1.0L) NXR_0002
Xla.Tg(LCMV:eYFPeCFP)Ryff NXR_0003
Wild Type Frog Stocks
Research Facility Resource
Workshops and Meetings
Back
Workshops and Meetings - Overview
XRET Program
Contact The NXR
Diving at MBL
Back
Diving at MBL - Overview
Dive Manual
10X Genomics Chromium Controller
Live Organism Use Form
Research Administration
Research Administration
Back
Research Administration - Overview
Contact Research Staff
Office of Sponsored Programs
Back
Office of Sponsored Programs - Overview
Sponsored Programs
Back
Sponsored Programs - Overview
Types of Agreements & Roles
Gifts vs. Grants
Uniform Guidance Information
Back
Uniform Guidance Information - Overview
Federal Agency Guidelines and Procurement
Subawards
Costs and Fees
Admin Salaries and Conferences
PI Absence from Project
Closeout
Grant & Contract Administration
Back
Grant & Contract Administration - Overview
Conflict of Interest and PHS (Public Health Science) Sponsored Specific Requirements
Effort Reporting: Semiannual Payroll Verification
Federal Policies & OSP Procedures
Funding & Sponsor Information
NIH and NSF Registration Requests
NSF Off-Campus/Off-Site Research (SAIF)
OSP Forms
All OSP News
Back
All OSP News - Overview
Announcing a Simplified Review Framework for NIH Research Project Grant Applications
NIH Grants Family-Friendly Initiatives
NIH Simplified Peer Review Framework
NIH: Preparing for 2025 Application and Review Changes – Tips and Resources
NIH’s Adoption of Common Forms for Biographical Sketch and Current and Pending (Other) Support by May 25, 2025
Revision: Notice of Updated Effective Date for the 2024 NIH Public Access Policy
Single Audit Requirements for Foreign Recipients and Subrecipients of NIH Grants
Updated Guidance on Salary Limitation for Grants and Cooperative Agreements
Updated NIH Policy on Foreign Subawards
Updated NIH Processes for No-Cost Extensions
eRA Information: New Data Management and Sharing (DMS) Policy Questions in RPPR as of October 1, 2024, and Other DMS Changes
Behavioral Codes of Conduct for NIH Award Recipients
Issuance of the NSF Proposal and Award Policies and Procedures Guide (PAPPG) (NSF 24-1)
NIH Application Instruction Updates – Data Management and Sharing (DMS) Costs
NIH Guidance on Marking Changes in Resubmission Applications
NSF Project Report Enhancements for Datasets and Research Materials
NSF-PAR System Updates Including Two Additional Product Types
Overview of NIH Grant Application and Review Changes for Due Dates on or after January 25, 2025
Reporting Data Management and Sharing (DMS) Plan Activities in the Research Performance Progress Report (RPPR)
Revised NSF Biographical Sketch and Current and Pending (Other) Support Formats - Now Available in SciENcv and on the NSF Website
Revision of NSF Award Terms and Conditions Applicable On or After May 20, 2024
Salary Cap Guidance Effective January 1, 2024
Updated Information about the Data Management and Sharing Plan from the NSF Directorate of Biological Sciences
Contact OSP
For Foreign Nationals
Back
For Foreign Nationals - Overview
Exchange Visitors Program
Back
Exchange Visitors Program - Overview
J-1 Visa Instructions
English Proficiency Policy for International Participants
Resources
Tax Information for Recipients of Scholarships, Fellowships, Stipends and Honoraria
Back
Tax Information for Recipients of Scholarships, Fellowships, Stipends and Honoraria - Overview
Applying for SSN or Individual Taxpayer ID
Foreign National Information System (FNIS)
Visas, Employment Restrictions, and Payments
Types of Visas
Contact Foreign Nationals Staff
Compliance and Ethics
Back
Compliance and Ethics - Overview
IACUC
Back
IACUC - Overview
IACUC Charter
Reporting Animal Care Concerns
Committee and Regulations
Animal Use Guidelines and Purchasing
Independent Review Board
Intellectual Property
Policies
Resources
Training
Contact HR Director
MBL Toolbox
Research Registration Forms
Funding Opportunities
Back
Funding Opportunities - Overview
Whitman Center Fellowships
Back
Whitman Center Fellowships - Overview
Whitman Center Fellowships Supporting Funds
Whitman Center Research Award Recipients
Whitman Center Research Awards FAQ
Grass Foundation
Health & Safety
Health & Safety
Back
Health & Safety - Overview
Biological Safety
Back
Biological Safety - Overview
Biosafety Committee
IBC Application
IBC Charter
Chemical Safety
Back
Chemical Safety - Overview
Acutely Hazardous Chemicals
Chemical Inventory Instructions
Chemical Storage and Segregation
Peroxide Forming Chemicals (PFC) Guidelines
Safety Data Sheet (SDS) Information
DEA Controlled Substances
Occupational Health and Safety
Back
Occupational Health and Safety - Overview
Electrical Safety
Fall Protection Program
Fire Safety Policy
Forklift Safety
Hearing Conservation
Ladder Safety
Lockout/Tagout Program
Machine Guarding
Occupational Health Program for Animal Handlers
Occupational Health and Safety Program
Workstation Ergonomics
Radiation and Laser Safety
Back
Radiation and Laser Safety - Overview
Radiation Safety Committee Charter
Radiation Safety Committee
Research Safety Questionnaire Form
Waste Management
Contact EHS Staff
Education
Back
Educational Opportunities
Advanced Research Training Courses
Advanced Research Training Courses
Back
Advanced Research Training Courses - Overview
Course Offerings
Back
Course Offerings - Overview
Analytical and Quantitative Light Microscopy
Back
Analytical and Quantitative Light Microscopy - Overview
Course Faculty and Lecturers
Financial and Course Support
2025 AQLM Schedule
Biology of Aging
Back
Biology of Aging - Overview
Course Faculty and Lecturers
Financial and Course Support
2023 Biology of Aging Schedule
Biology of Parasitism: Modern Approaches
Back
Biology of Parasitism: Modern Approaches - Overview
Course Faculty and Lecturers
Financial and Course Support
2024 BoP Schedule
Student Testimonials
Blout Lectureship
Englund Lectureship
Biology of the Inner Ear: Experimental and Analytical Approaches
Back